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How Green Was My Valley

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I watched "How Green Was My Valley" yesterday and noticed that one of the older brothers, I'm sorry to say I couldn't understand what they called him, but he was prominently played, looks EXACTLY like Nathan Fillion who plays Richard Castle in "Castle" on TV. They could be twins. I wondered if there is any relation. I've searched IMDB, etc, but no luck in identifying the actor. He's very obvious in the film, with many scenes.

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